International Journal of Innovative Approaches in Agricultural Research
Abbreviation: IJIAAR | ISSN (Online): 2602-4772 | DOI: 10.29329/ijiaar

Original article    |    Open Access
International Journal of Innovative Approaches in Agricultural Research 2019, Vol. 3(1) 25-52

Estimation of Gene Action, Heritability and Pattern of Association among Different Yield Related Traits in Upland Cotton

Bilal Nawaz, Muhammad Naeem, Tanwir Ahmad Malik, Ghulam Muhae-Ud-Din, Qadeer Ahmad & Saira Sattar

pp. 25 - 52   |  DOI:

Published online: March 29, 2019  |   Number of Views: 365  |  Number of Download: 854


A random mating design comprising 6 varieties (FH-142, MS-DK, IUB-63, KZ-189, FB-3159, and VH-300) and F1 crosses was used to check gene action for yield contributing traits in upland cotton.  Heritability and correlation were calculated from F3 populations of the cross CRS-456×Jumbo okra. The randomized complete block design was used in both experiments with three replication at the Department of Plant Breeding and Genetics, University of Agriculture Faisalabad, Pakistan during 2015-2017. In both experiments, analysis of variance indicated significant variation present among the accessions of the upland cotton for all the traits (plant height, sympodial and monopodial branches, nodes number of 1st sympodial branch, height of 1st sympodial branch, bolls per plant, boll weight, lint weight, seed cotton yield, seed index, lint percentage, lint index, seeds per boll, fiber fineness, fiber strength, fiber length and fiber uniformity) under study. Mean values of all genotypes were significantly different from each other for all the traits studied. All the parameters manifested positive correlation with seed cotton yield except plant height, monopodia per plant, nodes number of 1st sympodia, height of 1st sympodia, bolls per plant, seed index and lint index. The inheritance of all the traits in F1 crosses was generally controlled by overdominance gene action except monopodial branches, nodes number of 1st sympodial branch, height of 1st sympodial branches, boll weight, fiber length and fiber uniformity that were partially controlled under additive gene action. Epistasis was not found to be involved in any of the traits. Estimation of broad sense heritability (h2 b. s) in F3 populations were high (60-97%) for all the traits under study. Results suggested form heritability and correlation that these traits can be improved either through appropriate selection method or hybrid breeding programme.

Keywords: Correlation, Heritability, Gene action, Yield related traits, Upland cotton (Gossypium hirsutum. L).

How to Cite this Article

APA 6th edition
Nawaz, B., Naeem, M., Malik, T.A., Muhae-Ud-Din, G., Ahmad, Q. & Sattar, S. (2019). Estimation of Gene Action, Heritability and Pattern of Association among Different Yield Related Traits in Upland Cotton . International Journal of Innovative Approaches in Agricultural Research, 3(1), 25-52. doi: 10.29329/ijiaar.2019.188.4

Nawaz, B., Naeem, M., Malik, T., Muhae-Ud-Din, G., Ahmad, Q. and Sattar, S. (2019). Estimation of Gene Action, Heritability and Pattern of Association among Different Yield Related Traits in Upland Cotton . International Journal of Innovative Approaches in Agricultural Research, 3(1), pp. 25-52.

Chicago 16th edition
Nawaz, Bilal, Muhammad Naeem, Tanwir Ahmad Malik, Ghulam Muhae-Ud-Din, Qadeer Ahmad and Saira Sattar (2019). "Estimation of Gene Action, Heritability and Pattern of Association among Different Yield Related Traits in Upland Cotton ". International Journal of Innovative Approaches in Agricultural Research 3 (1):25-52. doi:10.29329/ijiaar.2019.188.4.

  1. Abbas, A., M. A. Ali and T.M. Khan (2008). Studies on gene effects of seed cotton yield and its attributes in five American cotton cultivars. J. Agric. Soc. Sci., 4, 147-152. [Google Scholar]
  2. Abbas, H.G., A. Mahmood and Q. Ali (2013). Genetic variability, heritability, genetic advance and correlation study in cotton (Gossypium   hirsutum L.). Int. Res. J. Microbiol., 4, 156-161. [Google Scholar]
  3. Ahmad, L., A. Ali, M. Zubair and I.A. Khan (2001). Mode of gene action controlling seed cotton yield and various components in (Gossypium hirsutum L.). Pak. J. Agri. Sci., 38, 19-21. [Google Scholar]
  4. Ahmad, M., N.U. Khan, F. Mohammad, S.A. Khan, I. Munir, Z. Bibi and S. Shaheen (2011). Genetic potential and heritability study for some polygenic traits in cotton (Gossypium hirsutum L.).  Pak. J. Bot., 43, 1713-1718. [Google Scholar]
  5. Ahsan, M.Z., M.S. Majidano, H. Bhutto, A.W. Soomro, F.H. Panhwar, A.R. Channa and K.B. Sial (2015). Genetic variability, coefficient of variance, heritability and genetic advance of some (Gossypium hirsutum L.) accessions. J. Agric. Sci. 7(2), 9752-9760. [Google Scholar]
  6. Akbar, M., J. Ahmad and F.M. Azhar (1994). Genetic correlation, path coefficient and heritability estimates of some important traits in upland cotton. Pak. J. Agric. Sci., 31(1), 47-50. [Google Scholar]
  7. Ali, M.A., A. Abbas, M. Younas, T.M. Khan and H.M. Hassan (2009). Genetic basis of some quantitative traits in upland cotton (Gossypium hirsutm L.). Plant Omics J., 2(2), 91-97. [Google Scholar]
  8. Ali. M.A., M.F. Bhatti. A. Abbas and I.A. Khan (2010). Assessment of inheritance pattern of some multigenic characters in cotton (Gossypium hirsutum L.). J. Agric. Res., 48, 1. [Google Scholar]
  9. Alkuddsi, Y., S.S. Patil, S.M. Manjula and B.C. Patil (2013). Correlation studies on yield and its components in interspecific cotton hybrids (G.hirsutum× G.barbadense) for developing heterotic box. Mol. Plant Breed., 4(28), 228-237. [Google Scholar]
  10. Altaher, A.F. and R.P. Singh (2003). Yield components analysis in upland cotton (Gossypium hirsutum L.). J. Ind. Soc. Cotton Improv., 28, 151-157.  [Google Scholar]
  11. Anapurve, S.N., G.S. Desmukh, S.P. Kausale, S.P. and U.V. Kale (2007). Correlation study of yield contributing characters in American cotton. J. Cotton Res. Dev., 21, 16-20. [Google Scholar]
  12. Anonymous (2015-16). Govt. of Pakistan, Ministry of Finance, Economic Advisor’s Wing, Islamabad. [Google Scholar]
  13. Arshad, M., M. Hanif, I. Noor, S.M. Shah and N. Hani (1993). Correlation studies on some commercial cotton varieties of Gossypium hirsutum L. Sarhad J. Agric., 9, 49-53. [Google Scholar]
  14. Ashokkumar, K. and R. Ravikesavan (2010). Genetic study of correlation and path coefficient analysis for seed oil, yield and fibre quality traits in cotton (Gossypium hirsutum L.). Aust. J. Basic Appl. Sci., 4, 5496-5499. [Google Scholar]
  15. Asif, M., J.I Mirza and Y. Zafar (2008). Genetics analysis for fibre quality of some cotton genotypes. Pak. J. Bot., 40, 1209-1215. [Google Scholar]
  16. Azhar, F.M. and S.U.K. Ajmal (1999). Diallel analysis of oil content in seed of Gossypium hirsutum L. J. Genet. Breed., 53, 19-23. [Google Scholar]
  17. Azhar, F.M., M. Naveed and A. Ali (2004). Correlation analysis of seed cotton yield with fiber characteristics in Gossypium hirsutum L. Int. J. Agric. Biol., 6, 656-658. [Google Scholar]
  18. Baloch, A., M.W. Baloch, M.K. Baloch, I.A. Mallano, A.M. Baloch, N.J. Baloch and S. Abro (2015). Association and Heritability Analysis for Yield and Fiber Traits in Promising Genotypes of Cotton (Gossypium hirsutum L.). Sindh Univ. Res. J., 47(2), 303-306. [Google Scholar]
  19. Baloch, M.J. and N.F. Veesar (2007). Identification of plant traits for characterization of early maturing upland cotton varieties. Pak. J. Sci. & Ind. Res., 50, 128-132. [Google Scholar]
  20. Baloch. M. J., C. Kumar, W.A. Jatoi and I.H. Rind (2014). Phenotypic correlation and regression analysis of yield and fiber quality traits in upland cotton (Gossypium hirsutum L.). Pak. J. Agri. Engg. Vet. Sci., 30, 135-146. [Google Scholar]
  21. Batool, S.N,U. Khan, K. Makhdom, Z. Bibi, G. Hassan, K.B. Marwat, Farhatullah, F. Mohammad, Raziuddin and I.A. Khan (2010). Heritability and genetic potential of upland cotton genotypes for morpho-yeild traits. Pak. J. Bot., 42, 1057-1064. [Google Scholar]
  22. Berdymuradov, R., R. Meredov and G. Novruzov (1990). Significance of backcrosses. Khlopok, 5, 42-43. [Google Scholar]
  23. Bibi M., N.U. Khan, F. Mohammad, R. Gul, A.Z. Khakwani and O.U. Sayal (2011). Genetic divergence and association among polygenic characters in Gossypium hirsutum L. Pak. J. Bot., 43(6), 2751-2758. [Google Scholar]
  24. Bing, T., J.N. Jenskins, C.E. Watson, J.C. McCarty and R.G. Grecer (1996). Evaluation of genetic variances and correlation for yield and fibre traits among cotton F1 hybrid populations. Euphytica, 91, 315-322. [Google Scholar]
  25. Brubaker, C.L., F.M. Bourland and J.E. Wendel (1999a).The origin and domestication of cotton. In: Smith, C.W. and J.T. Cothren, (Eds.), Cotton: Origin, History, Technology and Production. John Wiley and Sons, Inc., New York: 3-31. [Google Scholar]
  26. Butany, W.Y., A.D. Munshi and A.S. Chopra (1966). Interrelationship between some characters in Gossypium hirsutum L. Ind. J. Genet., 26, 262-268. [Google Scholar]
  27. Chao-zhu, X., Y. Shun-xun, G. Li-Ping, M. Cheng-duo, F. Wen-Juan, W. Hai-Lin and Z. Yun-Lei. (2007).  Heterosis performance and correlation analysis on economic traits of upland cotton in different ecological environments. Cotton Sci., 19, 3-7. [Google Scholar]
  28. Cherry, J.P. and Loffler, H.R. 1984. Seed. In: Kohel, R.J. and Lewis, C.F. (eds.), Cotton, ASA, Madison, WI: 511-569.  [Google Scholar]
  29. Desalegn D., N. Ratanadilok and R. Kaveeta (2009). Correlation and heritability for yield and fiber quality parameters of ethiopian cotton (Gossypium Hirsutum L.) estimated from 15 (diallel) crosses. Kasetsart J., 43, 1–11. [Google Scholar]
  30. Ekinci, R., S. Basbag and O. Gencer (2010). Path coefficient analysis between seed cotton yield and some characters in cotton (Gossypium hirsutum L.). J. Envir. Biol., 31, 861-864. [Google Scholar]
  31. Erande, C.S., H.V. Kalpande, D.B. Deosarkar, S.K. Chavan, V.S. Patil, J.D. Deshmukh, V.N. Chinchane, A. Kumar, U. Dey and M.R. Puttawar. 2014. Genetic variability, correlation and path analysis among different traits in desi cotton (Gossypium arboreum L.). Afric. J. Agri. Res., 9, 2278-2286. [Google Scholar]
  32. Farooq, J., M. Anwar, M. Riaz, A. Farooq, A. Mahmood, M.T.H. Shahid, M. Rafiq, S. and F. Ilahi  (2014). Correlation and path coefficient analysis of earliness, fiber quality, and yield contributing traits in cotton (Gossypium hirsutum L.). J. Animal Plant Sci., 24(3), 781-790. [Google Scholar]
  33. Farooq, J., M. Anwar, M. Riaz, A. Mahmood, A. Farooq, M.S. Iqbal and M.S. Iqbal (2013). Association and path analysis of earliness, yield and fiber related traits under cotton leaf curl virus intensive condition in Gossypium hirsutum L. Plant Know. J., 2, 43-50. [Google Scholar]
  34. Fryxell, P.A. (1979). The natural history of the cotton tribe. Texas A & M University Press, College Station, Texas, USA. [Google Scholar]
  35. Giri, A.N. and U.C. Updhyay (1980). Correlation and regression studies in upland cotton under different patterns and intercropping systems. Inf. J. Agric. Sci., 50, 907-910. [Google Scholar]
  36. Hendawy, F.A., M.S. Rady, A.M. Hamid and R.M. Esmail (1999). Inheritance of fibre traits in some cotton crosses. Egyptian J. Agron., 21,  35-36. [Google Scholar]
  37. Hussain, A., F.M. Azhar, M.A. Ali, S. Ahmad, K. Mahmood (2010a). Genetic studies of fiber characters in upland cotton. Animal Plant Sci., 20(4), 234-238. [Google Scholar]
  38. Hussain, S., N.N. Nawab, M.A. Ali, A. Hussain, M.A. Nawaz and T.A. Malik (2010). Evolution of performance, genetic divergence and character association of some polygenic traits in upland cotton. J. Agric. Soc. Sci., 6(4), 79-82. [Google Scholar]
  39. Hussain, S.B., Z. Ali and H.N. Khan (2014). A-one: high yielding, CLCV tolerant and transgenic Bt cotton variety for irrigated areas of Pakistan. J. Ann. Plant Sci., 24, 543-549. [Google Scholar]
  40. Imran, M., A. Shakeel, J. Farooq, A. Saeed, A. Farooq and M. Riaz (2011). Genetic studies of fiber quality parameter and earliness related traits in upland cotton (Gossypium hirsutum L.). AAB. J. Bioflux, 3, 151-159. [Google Scholar]
  41. Iqbal, M., K. Hayat, R.S.A. Khan, A. Sadiq and Noor-ul-Islam (2006). Correlation and path coefficient analysis for earliness and yield traits in cotton (Gossypium hirsutum L.). Asian J. Plant Sci. 5(2), 341-344. [Google Scholar]
  42. Iqbal, M., M.A. Chang, M.Z. Iqbal, M.U. Hassan, A. Nasir and N.U. Islam (2003). Correlation and path coefficient analysis of earliness and agronomic characters of upland cotton in Multan. Pak. J. Agron., 2(3), 160-168. [Google Scholar]
  43. Iqbal, M.Z. and M.A. Nadeem (2003). Behavior of some polygenic characters in cotton (Gossypium hirsutum L.). Asian. J. Plant Sci., 2, 485-490. [Google Scholar]
  44. Iqbal. M, M.A. Khan, M. Jameel, M.M. Yar, Q. Javaid, M.T. Aslam, B. Iqbal, S. Shakir, A. Ali (2011). Study of heritable variation and yield components in upland cotton. (Gossypium hirsutum L.). Afr. J. Agric. Res., 6, 4099-4103. [Google Scholar]
  45. Jatt, T., H. Abro, A.S. Larik and Z.A. Soomro (2007). Performance of different cotton varieties under the climatic conditions of Jamshoro. Pak. J. Bot., 39, 2427-2430. [Google Scholar]
  46. Joshi, H.J., P.K. Chovatia and D.R. Mehta (2006). Genetic variability, character association and component analysis in upland cotton. Ind. J. Agric. Res., 40, 302-305. [Google Scholar]
  47. Kalim-Ullah, Z. Usman, N. Khan, Rehmant-Ullah, F.Z. Saleem, S.I. Khattak and M. Ali (2015). Genetic diversity for yield and related traits in upland cotton genotypes. Pak. J. Agric. Res., 28, 118-125. [Google Scholar]
  48. Kapoor, C.J. and S.K. Kaushik (2003). Variability, heritability and genetic advance studies in cotton (Gossypium hirsutum L.). J. Cotton Res. Dev., 17, 242. [Google Scholar]
  49. Karademir, E., C. Karademir, R. Ekinci and O. Gencer (2010). Relationship between yield, fiber length and other fiber related traits in advanced cotton strains. Not. Bot. Hort. Agrobot. Cluj., 38(3), 111-116. [Google Scholar]
  50. Khan, A.I. and F.M. Azhar (2000). Estimates of heritabilities and pattern of association among different characters of Gossypium hirsutum L. Pak. J. Agri., 37, 7-10. [Google Scholar]
  51. Khan, M.I., M.A. Dasti, Z. Mahmood, M.S. Iqbal (2014). Effects of fiber traits on seed cotton yield of cotton (Gossypium hirsutum L.). J. Agric. Res. 52(2), 159-166. [Google Scholar]
  52. Khan, N.U. (2003). Genetic analysis, combining ability and heterotic studies for yield, its components, fibre and oil quality traits in upland cotton (G. hirsutum L.). Ph. D Dissert. Sindh Agric. Univ. Tandojam, Pakistan. [Google Scholar]
  53. Khan, N.U., H.K. Abro, M.B. Kumbhar, G. Hassan and M. Khan (1999). Exploitation of heterosis can combat Cotton Leaf Curl Virus (CLCV) incidence in cotton (G. hirsutum L.). The Pak. Cottons., 43(3&4), 21-33. [Google Scholar]
  54. Khan, N.U., K.B. Marwat, G. Hasssan, M.B. Kumbhar, Farhatullah, Z.A. Soomro, N. Khan, A. Parveen and U. Aiaman (2009). Study of fiber quality traits in upland cotton using additive-dominance model. Pak. J. Bot., 41, 1271-1283. [Google Scholar]
  55. Killi, F., L. Efe and S. Mustafayev (2005). Genetic and environmental variability in yield, yield components and lint quality traits of cotton. Int. J. Agric. Biol., 7, 1007-1010. [Google Scholar]
  56. Larik, A.S., A.A. Kakar, M.A. Naz and M.A. Shaikh (1999). Character correlations and path analysis in seed cotton yield (G.  hirsutum L.). Sarahad J. Agric., 15, 269-274. [Google Scholar]
  57. Lu, H. and G.O. Myers (2011). Combining abilities and inheritance of influential upland cotton varieties. Aust. J. Crop Sci., 5, 384-390. [Google Scholar]
  58. Ma, r.c., Y.Y. Zhou, R.T. Wang and C.O. Liu (1983). Genetic analysis of characters in the progeny of an intervarietal hybrid of upland cotton. Acta Agri. Univ. Pekinansis (China) 9(4), 27-34 PI. Br. Abst. 54(4-5), 3321-1984. [Google Scholar]
  59. Magadum, S., U. Baneijee, R. Ravikesavan, K. Thiyagu, N.M. Boopathi and S. Rajarathinam (2012). Association Analysis of Yield and Fibre Quality Characters in interspecific Population of Cotton (Gossypium spp.). J. Crop Sci. Biotech., 15, 239-243. [Google Scholar]
  60. Majeedano, M.S., M.Z. Ahsaan, A.W. Somroo and A.R. Channa (2014). Heritability and correlation estimates for some yield traits of Gossypium hirsutum.  Am. Res. Thoughts, 1, 781-790. [Google Scholar]
  61. Malagouda, P., B.M. Khadi, K. Basamma and I.S. Katageri (2014). Genetic variability and correlation analysis for fiber quality traits in diploid cotton (Gossypium spp). American-Eurasian J. Agric. Environ. Sci., 14(5), 392-395. [Google Scholar]
  62. Meena, R.A., D. Monga and R. Kumar (2007). Undescriptive cotton cultivars of north zone: an evaluation. J. Cotton Res. Dev., 21(1), 21-23. [Google Scholar]
  63. Mei, Y., W. Hu, S. Fan, M. Song, C. Pang and S. Yu (2013). Analysis of decision-making coefficient of three main fibre quality traits for upland cotton (Gossypium hirsutum L.). Euphytica, 140, 946-961. [Google Scholar]
  64. Murtaza, N. (2005). Study of gene effects for boll number, boll weight, and seed index in cotton. J. Central Eur. Agric., 6, 255-262. [Google Scholar]
  65. Mustafa, A.M., Y.M.A. Elsheikh and E.A. Babikar (2007). Genetic variability and character association and selection criteria in cotton (Gossypium hirsutum L.). Sudan J. Agric. Res., 8, 43-50. [Google Scholar]
  66. Nachnani, G.H. and H.K. Abro (1980). Correlation studies of yield with certain physical characteristics in F3 generations of cross of Upland cotton. The Pak. Cottons, 24, 119-129. [Google Scholar]
  67. Natera, J.R.M., A. Rondon, J. Hernandez and J.F.M. Pinto (2012). Genetic studies in upland cotton for genetic parameters, correlation and path analysis. SABRAO J. Breed. Genet., 44, 112-128. [Google Scholar]
  68. Naveed, M., F.M. Azhar and A. Ali (2004). Estimates of heritabilities and correlations among seed cotton yield and its components in Gossypium hirsutum L. Int. J. Agric. Biol., 6, 712-714. [Google Scholar]
  69. Ping, L.J., M.Y. Jun, Z.L. Li, H.S. Lin, G.W. Feng and X.R. Cinpek (2005). Analysis of heredity and correlation between boll traits and fibre quality traits in "0" plant type island cotton. Acta Agronomica Sinica, 31, 1069-1073. [Google Scholar]
  70. Poehlman, J.M. and D. Sleeper (1995). Breeding field crops. 4th ed. Iowa State University Press, Ames, Iowa: 369-370. [Google Scholar]
  71. Pujer, S., S.S.J. Deshmukh, R.S. Sangwan and O. Sangwan (2014). Genetic variability, correlation and path analysis in upland cotton (Gossypium hirsutum L.). Electron. J. Plant Breed., 5, 284-289. [Google Scholar]
  72. Rajarathinam, S., N. Nadarajan, Sukanya and Subramaniam (1993). Genetic variability and association analysis in cotton. J. Ind. Soc. Cotton Improv., 18, 54-59. [Google Scholar]
  73. Ranganatha, H.M., S.S. Patill, S.M. Manjula and B.N. Avinkumar (2011). Genetic variability studies in segregating generation of upland cotton (Gossypium hirsutum L.). Mol. Plant Breed., 4, 84-88. [Google Scholar]
  74. Rasheed, A.,W. Malik, A.A. Khan, N. Murtaza, A. Qayyum and E. Noor (2009). Genetic evaluation of fiber yield and yield components in fifteen cotton (Gossypium hirsutum) genotypes. Int.  Agri. Biol., 11(5), 581-585. [Google Scholar]
  75. Reddy, Y.R. and A.S.R. Sharma (2014). Genetic variability for yield components and fiber characters in cotton (Gossypium hirsutum L.). Plant Arch., 14, 417-419. [Google Scholar]
  76. Saghir, A., M.Z. Iqbal, A. Hussain, M.A. Sadiq and A. Jabbar (2003). Gene action and heritability studies in cotton (Gossypium hirsutum L.). Pak. J. Biol. Sci., 3, 443-450. [Google Scholar]
  77. Salahuddin, S., S. Abro, M.M. Kandhro, L. Salahuddin and S. Laghari (2010). Correlation and path analysis of yield component of upland cotton (Gossypium hirsutum L.).World Appl. Sci. J., 8, 71-75. [Google Scholar]
  78. Salahuddin, S., S. Abro. A. Rehman and K. Iqbal (2010). Correlation analysis of seed cotton yield with some quantitative traits in upland cotton (Gossypium hirsutum L.). Pak. J. Bot., 42, 3799-3805. [Google Scholar]
  79. Sarwar, S., M. Baber, N. Hussain, I.A. Khan, M. Naeem, M.A. Ullah and A.A. Khan (2011). Genetic dissection of yield and its components in upland cotton (Gossypium hirsutum L.). Afr. J. Agri. Res., 6, 2527-2531. [Google Scholar]
  80. Selim, A.K.A., A.M. Omar, A.M. El-Marakby and M.A. Mohamed (1978). Inheritance of lint percentage and its correlation with fibre strength in a cross between two varieties of Egyptian cotton. Aim. Shams Univ. Agric. Faculty. Res. Bull., 948. [Google Scholar]
  81. Shakeel, A., I. Talib, M. Rashid, A. Saeed, K. Ziaf and M.F. Saleem (2015). Genetic diversity among upland cotton genotypes for quality and yield related traits. Pak. J. Agric. Sci., 52, 73-77. [Google Scholar]
  82. Soomro, B.A., M.H. Channa and M. Ahmad (1982). Correlation studies in Gossypium hirsutum L. The Pak. Cottons, 26, 39-51. [Google Scholar]
  83. Soomro, Z.A., A.S. Larik, M.B. Kumbhar, N.U. Khan and N.A. Panhwar (2008). Correlation and path analysis in hybrid cotton. SABRAO. J. Breed. Genet., 40, 49-56. [Google Scholar]
  84. Soomro, Z.A., M.B. Kumbhar, A.S. Larik, M. Imran and S.A. Brohi (2010). Heritability and selection response in segregating generations of upland cotton. Pakistan J. Agri. Res., 23, 1-2. [Google Scholar]
  85. Suinaga, F. A., C.S. Bastos and L.E.P. Rangel (2006). Phenotypic adaptability and stability of cotton cultivars in Mato Grosso State, Brazil. Pesquisa Agropecuaria Trop. (PAT), 36(3), 145-150. [Google Scholar]
  86. Swati, B., L.D. Meshram and P.W. Khorgade (1999). Genetic variability and character association in naturally coloured cotton (Gossypium hirsutum L.). J. Ind. Soc. Cotton Improv., 24, 197-199. [Google Scholar]
  87. Tang, F. and W. Xio (2014). Genetic association of with in boll yield components and boll morphological traits with fibre properties in upland cotton (Gossypium hirsutum L.). Plant Breed., 133, 521-529. [Google Scholar]
  88. Taohua, Z. and Z. Haipeng (2006). Comparative study on yield and main agri-characters of five hybrids coloured cotton varieties. J. Anhui Agric. Univ., 33(4), 533-536. [Google Scholar]
  89. Thiyague, K., N. Nadarajan, S. Rajarathinam, D. Sudhakar and K. Rajendran (2010). Association and path analysis for seed cotton yield improvement in interspecific crosses of Cotton. Electron. J. Plant Breed., 1, 1001-1005. [Google Scholar]
  90. Tian, Z.G., X.Q. Haq and S.F. Zhang (1993). Inheritance analysis and selection strategies for earliness and yield component in short season cotton. Acta Agri. Borcali Sinica., 8, 18-22. [Google Scholar]
  91. Turner, J.H., H.H. Ramey and S. Workley (1976). Influence of environment on seed quality of four cotton cultivars. Crop Sci., 16, 407-409. [Google Scholar]
  92. Vineela, N., J.S.V.S. Murthy, P.V. Ramakumar and S.R. Kumari (2013). Variability studies for morphological and yield components traits in American cotton (Gossypium hirsutum L.). J. Agric. Vet. Sci., 4, 7-10. [Google Scholar]
  93. Waldia, R.S., D.S. Jatasra and B.N. Dahiya (1979). Correlation and Path analysis of yield components in Gossypium arboreum L. Ind. J. Agri. Sci., 49, 32-34. [Google Scholar]
  94. Wang, C., A. Isoda and P. Wang (2004). Growth and yield Performance of some cotton cultivars in Xinjiang, China, an arid area with short growing period. J. Agron. Crop Sci., 190 (3):177–183. [Google Scholar]
  95. Wu, J., J.N. Jinkins, J.C. McCarty and J. Zhu (2004). Genetic association of yield with its components in recombinant inbred line population of cotton. Euphytica, 140, 171-179. [Google Scholar]
  96. Yuan, Y.L., T.Z. Zhang, W.Z. Guo, J.J. Pan, Y.L. Yuan, T.Z. Zhang, W.Z. Guo and J.J. Pan (2002). Genetic stability of fibre qualities in Upland cotton. Cotton Sci., 14, 67-70. [Google Scholar]
  97. Zeng, L., W.R. Meredith, L. Deborah, Boykin and E. Taliercio (2007). Evaluation of an exotic germplasm population derived from multiple cross among Gossypium tetraploid species. J. Cotton Sci., 11, 118-127. [Google Scholar]